I had a look at the schematic for the 180A, and the neon bulbs are used as voltage limiters between the CRT cathode and control grid (g1).
If they light up during normal operation, then I'd expect if to be fairly weakly. When they flash, then it is because you have what you describe: A spark jumping on either the -2.9KV or the slightly less negative control grid rail (supplied by CR302), temporarily causing a huge voltage difference across the neon bulbs. Could be in any of the associated components, or maybe just due to dust buildup, as others have suggested.