In both the Blackbird and treadmill experiments the vehicle accelerated. Blackbird accelerated so vigorously they had to put the brakes on to slow it down. If an unpowered vehicle accelerates it is gaining energy from the surroundings. If it accelerates under its own steam, it will run forever, by definition.
Blackbird is build at the limit as they wanted to keep the weight down so that is the reason it looked it will self destroy as it was close due to vibrations.
The acceleration is from the stored energy and highest stored energy (full storage) is at around 0.7x the wind speed on blackbird and of course for the treadmill model highest acceleration is immediately after you release the vehicle from your hand.
You will be able to feel the force of the stored energy on your had as an elastic type force from the compressed (higher density) air behind the propeller.
As the model advances on treadmill stored energy will decrease and so will the acceleration rate but if you stop the vehicle by blogging it going forward with your hand the energy storage will be again filled up so as long as you keep the vehicle from moving you have just a treadmill powered propeller.
If there is a velocity difference between the air and the ground, then a vehicle in contact with both air and ground can extract energy from the system. This can happen even if the vehicle is moving faster than the differential velocity.
As far as vehicle is concerned when you are at exact same speed as wind speed (like in the treadmill example) air molecules do not move relative to the vehicle meaning air molecules can not transfer any energy to the vehicle. The only way vehicle accelerates from that point is due to stored energy as there is that high density air in the back of the propeller.
As vehicle accelerates past wind speed that pressure will slowly decrease. Slowly because most of the stored energy is put back in the propeller to maintain part of this pressure differential (that is what the connection between wheel and propeller will do).
At peak speed there will very little pressure differential between the front and back of the propeller, so small that it can no longer cover the vehicle losses and have extra to accelerate and that is the point where propeller and vehicle will have the highest speed and vehicle will start to slowly decelerate.
This part after peak speed is where the kinetic energy both the one in the vehicle body but also the propeller (flywheel) will start to be used thus the reason it will take probably as long to decelerate below wind speed as it took to get to peak speed.
If course wind is necessary! If there is no wind, there is no motive power for the vehicle, and it will coast to a standstill and stop moving.
If you push one of this propeller based vehicles either blackbird or the treadmill models (needs to be pushed to a minimum speed probably around 10mph maybe 11mph due to extra drag) then when released the vehicle will accelerate well past that speed before slowing down to zero and no wind at all is needed.
I think this is already shown in the treadmill example if you interpret that correctly as test is done in a room with no wind and so you keeping the vehicle on the treadmill is the same with pushing the vehicle to 10mph (small difference is the extra drag when pushed) and so when you let go while there is no wind you notice vehicle accelerates and it can not use energy from treadmill as vehicle advances on the treadmill thus that means it is not slowing down (or try to slow down) the treadmill. Meaning energy comes from the compressed air behind for lack of a better description.
This stored energy will eventually be transferred to vehicle in form of kinetic energy minus the losses trough friction. That means that if you where to add weight to the vehicle the vehicle will be able to travel less on the treadmill and if you add weight almost perfectly the vehicle will accelerate forward so slowly that you will be able to see how it slows down before it gets to the end of the treadmill.